Wednesday, March 19, 2014

Soil - properties and contamination

Soil is a living system that has to remain in a state of dynamic balance to be healthy. It is a complex ecosystem in which the substances move in cycles from plants to animals, to soil bacteria and back again to plants. Solar energy is the natural fuel that drives the soil cycles and living organisms of all kinds are necessary to sustain the whole system and keep it in balance. Soil is an essential base of human life.It is a  site for agricultural and forest production, a place for storing raw materials and wastes, a constituent element of landscape and a mirror of the history of civilizations and cultures.
Soil is a biologically active, complex mixture of minerals, organic materials, living organisms, air and water. Soil is an interface between biosphere, lithosphere, hydrosphere and atmosphere. Land occupies 38% of the total space. Soil is the outer most layer of the earth’s crust (Earth’s living skin-1/3 of surface). About 95% of human food is derived from the earth. Only 10% of the world’s land area is suitable for growing crops. Soil quality is degrading worldwide. Soil biota depend on the soil environment for their energy and nutrient supply.


 Soil functions 

Soil is an essential natural resource. It is an integrator of all parts of ecosystems. It is a medium for plant growth, a home for organisms and a storehouse of water, heat and chemicals.It is a decomposing medium for wastes, a source material for construction of shelter and a buffer system  to neutralize harsh environmental changes. Soil is a dynamic universal ecological system and a complex heterogeneous medium. A vital resource that provides food, feed, fuel and fiber. Soil is a material which nourishes and supports growing plants. Soil is a mixture of mineral matter, organic matter, water and air. (Example: Loam soil = 45% mineral matter, 5% organic matter, 25% water, and 25% air). Soil is a three phase system of a solid phase (Mineral matter,organic matter), a liquid phase (soil water) and a  gaseous phase( soil air).


Major components of soil

 Soils have four major components: Mineral matter contains three fractions, sand, silt, and clay. Organic matter contains appreciable quantities of nitrogen, phosphorus and sulfur. Air and water  occupy the pore spaces in soils.
Soil volume -Soil consists of organic particles and inorganic matter with pore spaces between and within them. Pore spaces contain soil air, and soil solution.  In other words, soil volume consists of solid, liquid and gaseous phases.

Types of soil water (i.e. water in the soil) 

 Gravitational water:  Gravitational water fills all the pore-space, and leaves no room for oxygen and gaseous exchange.
Capillary water: This water which is held with the force of surface tension by the soil particles, and is resistant to the forces of gravity. 
Hygroscopic water: This water is held so tightly (by surface tension) to the soil particles that the plant roots can't take it up.

Physical Properties of Soils 

 Soil Texture the relative proportions of sand, silt, and clay in the soil. e.g. sandy soil, silty soil,and clay soil.
Soil Structure – Structure refers to the arrangement of soil particles. E.g., Granular, Platy, Wedge, Blocky, Prismatic, and Columnar. Soil structure is of particular importance in the absorption of water and the circulation of air.
Soil Color - Color in soils is due primarily to two factors, humus content and the chemical nature of the iron compounds. Humus has a dark brown or black color. Iron is an important color material which stains mineral particles. Ferrous oxide gives gray color. Ferric oxide gives red color. Hydrated ferric oxide gives yellow color.
Soil pH is primarily controlled by the concentration of free hydrogen ions in the soil matrix. Acidic soils have a relatively large concentration of hydrogen ions. Alkaline soils have a relatively low concentration of hydrogen ions.
Soil Types - Depending on the size of the particles in the soil, it can be classified as:
sandy soil, silty soil, clay soil, loamy soil, peaty soil and chalky soil.

Soil contamination

It is the presence of man-made chemicals or other alteration to the natural soil environment. Contaminants bind tightly to the soil. Contaminants evaporate into the air and end up in soil/ground water. The common chemicals of soil contamination include petroleum hydrocarbons, pesticides, heavy metals and solvents.Sources of soil contaminants comprise industrial wastes, active mine wastes, solid wastes and waste waters.
 Petrochemicals- Consumption, transportation and extraction of fossil fuels. 
Agricultural chemicals- Pesticides, fertilizers.
Solid wastes - Wastes from agriculture – crop and farm residues, animal manure.
Wastes from mining – coal wastes, metal ore wastes.
Industrial wastes- solvents chemicals, paints.
Solids from sewage treatments -biomass sludge, settled solids.
Ashes – residues from solid fuels
Garbage – glass, metals, clothes, plastics, wood, papers.
The intensive use of agrochemicals reduces soil fertility, soil biodiversity, nitrogen fixation and crop yield. Pesticides may infiltrate soil, carried away by wind, spread by runoff, leach out into the groundwater and finally reach rivers, lakes and ocean.

Human health effects

Exposure to heavy metals in soil results in nervous system disorders, kidney damage and liver toxicity. Exposure to agricultural chemicals causes cancer and infertility. Chronic exposure to other industrial toxins may causes birth defects, nervous system disorders and kidney diseases.

Quote for reflection

"We abuse land because we regard it as a commodity belonging to us. When we see land as a community to which we belong, we may begin to use it with love and respect." 
-Aldo  Leopold, A sand county Almanac.

Tuesday, March 4, 2014

Endocrine system - characteristics and functions

Endocrinology is the science dealing with the chemical integration of the physiological functions of an organism.  The science of Endocrinology was born from the experiments of Bayliss and Starling(1902 to 1905). Pende introduced the term ‘Endocrinology’ (Greek, endon=within; krinein= to separate). There are two types of glands in human body: exocrine glands or glands with duct or glands of external secretion e.g. salivary glands and endocrine glands or ductless glands or glands of internal secretion e.g. pituitary gland, thyroid gland.
A hormone is a regulatory chemical that is secreted into the blood by an endocrine gland. The blood distributes the hormones throughout the body. The hormones produce specific physiological effect on certain target organs. The word hormone is derived from the Greek word ‘hormon’ which means ‘to excite’. Bayliss and Starling in 1905 coined the term ‘hormone’.


 Types of chemical messengers
·         Hormones- are products of endocrine glands.
·         Neuro-hormones – are products of nerve cells and released at neurohemal organ.
·         Neurohumors- are products of nerve cells and released at axons.
·         Phytohormones –plant hormones e.g.auxin.
·         Pheromones –ectohormones.

Properties of hormones

 Hormones are low molecular weight chemical messengers. They are secreted in trace amounts in response to specific secretary stimuli. They are soluble in water and act as catalysts.
A single hormone has multiple effects on a single target tissue or on several different tissues. Hormones have high degree of action specificity. They are not species specific and non-antigenic. They are inactivated as soon as their functions are over. Endocrine glands are under the control of nerves.

Chemical classes of hormones

1.     Amines - Secreted by gland cells of nervous origin. e.g.,Adrenaline, ADH.
2.     Steroids - Secreted by glands derived from coelomic mesothelium e.g., corticosteroids, estrogens, androgens.
3.     Proteins - Hormones made up of amino acids and polypeptides e.g., thyroxine, calcitonin, STH.
Steroid hormones are formed from cholesterol. They are lipid-soluble hydrophobic molecules. They are secreted by the gonads, adrenal cortex and placenta e.g., aldosterone, cotisol, estrogen, testosterone and progesterone.
Peptide hormones are short chains of amino acids. They are water-soluble. They are secreted by pituitary and parathyroid glands e.g., oxytocin, vasopressin, calcitonin, insulin
Amine hormones - They are derived from the amino acid tyrosine. They are secreted from the thyroid and the adrenal medulla.

Mechanism of hormone action

 Hormones activate or inhibit enzyme systems e.g., phosporylase. Hormones alter cell permeability. For example Insulin promotes transfer of glucose. Hormones directly activate or suppress particular genes. For example ecdysone causes puffing of certain genes.
Control of hormone secretion
 Humoral signal-secretion depends upon the level of blood ions and nutrients. E.g. insulin production controlled by blood glucose.
Neuronal signal- secretion depends on nerve impulses-stimulation of sympathetic nervous system release catecholamines.
Hormonal signal- some hormones control the release of hormones from another endocrine tissue-hypothalamus hormones.
Hormone secretion can be stimulated / inhibited by other hormones - e.g., trophic hormones, plasma concentrations of ions /nutrients, neurons and mental activity and environmental changes - e.g. light, temperature

Concept of feedback mechanism

The production of a hormone by an endocrine gland is controlled by its circulatory level. A reciprocal relationship exists between the blood level of a hormone and rate of its synthesis and secretion. Feedback mechanisms help in maintaining homeostasis within the endocrine system. This concept is proposed by Moore and Price in 1932. It is a two-way communication between the endocrine gland and the target gland. It is a self-balancing mechanism.
 In a negative feedback system, a gland is sensitive to the concentration of the substance it regulates. When the concentration of the regulated substance reaches a certain threshold level, it inhibits the gland from secreting more hormones until the concentration returns to normal. For example the neurons in the hypothalamus secrete thyroid releasing hormone (TRH), which stimulates the anterior pituitary to secrete thyroid-stimulating hormone (TSH). TSH stimulates the synthesis and secretion of thyroid hormones. When blood levels of thyroid hormones increase above a certain threshold, TRH-secreting neurons in the hypothalamus are inhibited and stop secreting TRH.
In a positive feedback, the increased activity of an endocrine gland is followed by stimulation. The positive feedback is not common. For instance oxytocin acts on uterine muscles during child birth. The secretion stops when baby leaves birth canal. The positive feedback is easily observed under experimental conditions e.g. muscular injection of estrogen in female induces ovulation.
Hormone transport 
Hormones are normally present in blood plasma. Steroid and thyroid hormones bind to transport proteins in the plasma. Amines and peptide hormones are hydrophilic and mix easily with blood plasma. Unbound hormones have shorter half life and bound hormones exhibit longer half life. Transport proteins protect circulating hormones.
Hormone receptors
Hormones activate only those cells that have receptors for them. Hormone receptors are located on plasma membrane, in the cytoplasm or in the nucleus. Hormone – receptor interactions exhibit specificity and saturation. Protein hormones react with receptors on the surface of the cell. Steroid hormones react with receptor sites inside a cell.

Functions of endocrine system

1.     Homeostatic function – endocrine system regulates body fluid composition, rate of gaseous exchange and cardiovascular functions.
2.     Integrative function – endocrine system supports the role of nervous system.
3.     Morphogenetic  function – It influences embryonic development
4.     Permissive function - Certain hormones require the presence of another hormone for the expression of their activity.

Endocrine disrupting chemicals (EDCs)

EDCs are both natural and man-made chemicals that may mimic or interfere with the function of hormones in the body. Environmental endocrine disrupting chemicals include persistent organic pollutants (POPs) such as pesticides (DDT), dioxins, polychlorinated biphenyls (PCBs) and plasticizers (biphenol A). Endocrine disruption is an important public health concern. EDCs produce adverse developmental, reproductive, neurological and immune effects in both humans and wildlife. In humans, EDCs may cause male infertility, reproductive abnormalities, reproductive diseases and neuro-developmental disorders.

Nerve impulse conduction -mechanism and characteristics

Nervous system consists of  interconnecting fibers of communication network. In the ‘hard-wiring’ of the nerves, the signals travel in the form of a flow of electrical current called nerve impulses. Irritability is the universal property of life which means the capacity of organisms to respond to changes in the environmental conditions. The specific environmental stimulus elicits a change in an organism is termed a response. The stimulus-response reactions afford internal constancy in the face of environmental changes.


Discovery of neuron
Camillo Golgi (1843-1926) invented a specific    staining technique for neurons. Cajal in 1888 identified the networks of nerve cells. Golgi and Cajal received the Nobel Prize in 1906 for Medicine and Physiology. Wilhelm His in 1886 showed that the dendrites and axons grow out progressively from the immature neurons in the brains of embryos. Henri Forel (1848-1931) in 1886,observed that when the  cell body dies or an axon is cut, degeneration  of the neuron stop at the junction to another neuron, thus giving evidence that they are separate.
Description of a neuron
A neuron consists of a cell body and two kinds of processes, the dendrites and the axon. The cell body has neuroplasm, a nucleus, nissl bodies, neurofibrils and a cell membrane. The dendrites carry impulses towards the cell body. The axon carries impulses away from the cell body. The axon originates from axon hillock of the cell body. The axon is surrounded by two coverings: myelin sheath and Schwann sheath. These two coverings are interrupted at intervals by nodes of Ranvier. The fine branches at the end of axon are called axon terminals.

Properties of neuron
1.    Excitability –a nerve can be stimulated by suitable stimulus-mechanical, thermal, chemical, electrical.
2.    Conductivity – impulse is conducted similar to cable conduction and digital in character.
3.    All or none law – the stimulus should be in adequate threshold strength.
4.    Refractory period – when the nerve fiber is once excited, it will not respond to a second stimulus for a brief period.
5.    Indefatigability – nerve is normally not fatigued.
6.    Adaptation –the nerve quickly adapts itself.
7.    Accommodation – slowly applied stimulus is accommodated.
Definition of nerve impulse
         A nerve impulse is the sum total of physical and chemical events associated with the transmission of a signal along an axon.
         A wave of physiological activity- primarily an electrical phenomenon.
Stimulus is defined as a sudden change in the environment which is strong enough to cause a response in the living organism. All or none law indicates the relation between the stimulus and response. A stimulus, if it is capable of causing a response, causes a maximum response. If it is below the capacity, it will not cause any response. The lowest strength of stimulus required to give rise to an action potential is the threshold stimulus. A stimulus which is less than the threshold fails to induce any response-sub-threshold stimulus. A stimulus which is greater than optimum is supra threshold stimulus.
Stimulation can be affected by strength and duration. The weaker the stimulus, the longer it will have to be applied to produce a response. The nerve takes lesser time to respond for a stronger stimulus.
Velocity of impulse conduction
Johannes Muller believed that nerve impulse travelled at a speed of light-186,000 miles per sec. Helmholtz showed that velocity of conduction was 100m per sec. (about 10 times faster than a man can run). The velocity varies from 100 m per sec in large fibers to 0.5 m per sec in small non-myelinated fibers.
Refractory period of impulse conduction
Once an impulse has passed over any part of the neuron, for a short time it is unable to conduct any other stimulus. This brief period of non-conductivity is called refractory period. Under good physiological conditions, the nerve fiber is indefatigable.
                   Mechanism of impulse conduction
Bioelectricity
L. Galvani in 1786 discovered the presence of electrical current in nerves and muscles.
Electrophysiology
Du Bois Raymond in 1848 concluded that impulse transmission was electrical by a wave of relative negativity.
Electrochemical basis of bioelectricity
Hodgkin and Huxley in 1939 demonstrated the electrical and chemical processes involved in bioelectricity.
Demonstration of electrogenesis – microelectrode studies
In order to confirm the generation of electricity, one microelectrode is placed on the outer surface of the nerve membrane and the other placed inside the nerve cell. When the terminals are connected to a galvanometer, the needle show a deflection indicating the flow of electrical current (-70mV). In a resting neuron the electrical potential ranges from 20 to 100 mV.

Resting potential

According to the membrane potential hypothesis of Bernstein (1902), the differential concentration of ions between the inside and outside of the nerve cell is the basis of resting potential. The inner side of the nerve contains large negatively charged non-diffusible protein ions and smaller diffusible K+ and Cl- ions. Na+ ions are more concentrated on outside of the nerve cell. Radio-isotopic studies showed that potassium and sodium ions readily diffuse through the nerve cell membrane.
In a state of physiological rest, the inner side of a neuron is electrically negative to outside. This difference in electrical charge is called resting potential. The resting potential is maintained as long as the cell is alive and active. The resting potential depends upon the selectivity and variable permeability of cell membranes. The unequal distribution of ions maintains the resting potential.
Factors influencing ionic imbalance in a nerve cell
·       Active transport
·       Concentration gradient
·       Membrane permeability
·       Electrostatic attraction

Ion-exchange pump - This Na+ - K+ pump maintains unequal concentration of ions in the nerve fibers by the process of active transport. This pump changes the electrical character of the nerve fibers. ATP is used as energy for the process.
Concentration gradient - The active transport establishes the concentration gradient. The extrusion of sodium ions is linked with active uptake of potassium ions. The rates of diffusion of ions depend upon gradients and membrane permeability.
Donnan membrane equilibrium - “at equilibrium, the product of the concentration of the diffusible ions inside the membrane equals the product of the concentration of the diffusible ions outside.” (Donnan 1928).
Selective membrane permeability - All the plasma membranes are selectively permeable to ions. This membrane selectively allows inward diffusion of K+ and prevents inward diffusion of Na+ ions.
Electrical gradient - The inner side of the nerve cell is negative to the outside. There is a growing attraction between ions of the same charges. The parallel diffusion of ions restores the resting potential.

Action potential (Hodgkin 1949, Huxley 1952 and Katz 1959)

When a nerve is stimulated, Na+ ions suddenly move into the cell and causing a positive potential. The influx of Na + ions reaches its peak in 100msec.Sodium permeability causes depolarization. The inward diffusion of Na + ions halted near the peak of action potential. The diffusion K+ ions restore the membrane potential –called repolarization. By using radioactive sodium (Na 24) and potassium (K42) Hodgkin and Keynes (1955) showed that inward flux of sodium was increased 20 times and outward flux of potassium 3-4 times after stimulation of nerve.



             Properties of Impulse conduction

The conduction of nerve impulse causes two phases in the moving wave called biphasic action potential. The conduction of nerve impulse in a myelinated fiber is so rapid because the action potential skips from node to node and this phenomenon is called saltatory conduction. The magnitude of nerve impulse transmission occurs without decrement is termed Non-decremental conduction. The velocity of conduction in a myelinated nerve is directly proportional to its diameter. The property of myelinization speeded up the conduction velocity and consumption of energy.

Muscle contraction - biochemical mechanisms and theories

A muscle contraction is a key biological process by which animals move by contracting their muscles. Movement is the basic property of living systems. The vital activities like digestion, reproduction, excretion and circulation are all possible by the contraction of muscles.
Muscles are complex biological motors, which convert chemical energy into mechanical work and force. In human beings, muscles constitute 40% of the body. The energy for muscle contraction is obtained from the chemicals adenosine triphosphate (ATP) and creatine phosphate ( CP).


                                          Uses of muscles

Skeletal muscle – skeletal muscle gives shape and structure to the body. It enables animals to maintain erect posture. It brings about movement. It helps the animal to secure food and shelter and escape from danger.It helps to communicate its wishes.
Smooth muscle - Smooth muscle assists breathing movements. It aids in hearing and vision.It helps the processes of digestion, excretion, reproduction and circulation. It helps to propel the digested food, body fluids, glandular secretions and waste products. It pumps blood to all parts of the body.
Characteristics of muscles
         Excitability - the ability to receive and respond to stimuli.
         Conductivity - The ability  to receive a stimulus and transmit a wave of excitation (electrochemical activity)
         Contractility - the ability to shorten forcibly when stimulated.
         Extensibility - the ability to be stretched or extended.
         Elasticity - The ability to bounce back to original length

                            Types of muscles

Unstriped or plain muscle – It has elongated spindle shaped muscle fibers with thickened central belly and two pointed terminals. The cytoplasm is granular without any cross striations. A rod like nucleus is placed in the center. Myofibrils are arranged in longitudinal axis. They are found in the alimentary canal, respiratory tract, uterus, urinary bladder, arteries and veins.
Cardiac or heart muscle – heart muscles are branched and form a net work. The two adjacent muscle cells form tight junctions in the form of intercalated discs.They exhibit faint transverse striations. They are innervated by autonomic nerve fibers. The nucleus is round or oval in shape. They have fewer myofibrils with greater amount of sarcoplasm for more storage of glycogen and the sarcolemma is indistinct.
Skeletal or striped or muscle – are complex, elongated, cylindrical and fast moving muscles. They vary considerably in size, shape and arrangements of fibers. The size ranges from the smallest stapedium muscle of the middle ear to larger thigh muscles of the human body. Each fiber is multinucleated with transverse and longitudinal striations. The cytoplasm is composed of myofibrils with many myofilaments. Large number tubules run through sarcoplasm and form sarcoplasmic reticulum. The sarcosomes or mitochondria supply ATP to the myofibrils.

        Light microscopic structure of skeletal muscle

Each muscle fiber  displays  dark anisotropic bands or  A bands and light  isotropic bands or I bands. Each A band has a less denser region called H band or Hensen’s line. In each I band, there is a dense cross line called Z band. The area between two adjacent Z lines is called a sarcomere.



Fibrillar system of skeletal muscle
The myofilaments consist of thick myosin filaments and thin actin filaments and are arranged in an overlapping manner. The myosin filaments bear thick knob like projections called cross bridges. The sarcoplasmic reticulum is made up of longitudinal system of canals between myofilaments. There is also T system of canals.

                          Chemical composition of muscle

Water - Muscle contains about 75-80% of water. Water provides a good medium for inorganic and organic compounds. Water reduces friction and dehydration of muscles during contraction.
Proteins - Muscle consists of 3 types of proteins namely structural proteins (e.g., collagen, elastin), contractile proteins (e.g., myosin, actin, and tropomyosin) and enzymatic proteins (e.g., adenosine triphosphatase, creatine phosphatase and lactic dehydrogenase).
Minerals - Calcium ions of sarcoplasm initiates muscle contraction. Magesium ions never initiate muscle contraction but important for muscle coordination. Sodium and Potassium ions set the action potential of impulse conduction.
Organic compounds - Muscle is a storehouse of glycogen and oxidation of glycogen provides energy.  The lipids found in the form of phospholipids. The activity of muscle is proportional to the amount of phospholipids. ATP is the primary source of energy for muscle contraction. ATP molecules found associated with G-actin.
Contractile proteins- Myosin is the  prime contractile element of muscle. It has a triple helical structure. Its molecular weight is 420,000. The hydrolysis of myosin with enzyme trypsin yields two fractions – heavy meromyosin (HMM) and light meromyosin (LMM). Hmm acts as an enzyme ATPase for splitting of ATP into ADP and Pi. The hydrolysis of HMM with papain yields sub- fragment 1 and sub-fragment 2.
Actin -  is non-contractile and elastic in nature. Actin is made up of spherical molecules (G-actin) with a molecular weight  of 60,000. G-actin polymerizes into double stranded helices called fibrous or F-actin.
G-actin +ATP -àF-actin +ADP +Pi.
           Association and disassociation of actomyosin
1 mole of actin +3 mole of myosin -à actomyosin (super- precipitation).
Actomyosin +ATP—ca++, mg++àactin +myosin +ADP
Tropomyosin – is a non-contractile, fibrous protein. It plays important role in sensitizing actin and myosin molecules to calcium ions. This sensitivity is important in order to switch contraction on or off.
Troponin - Troponin occurs at intervals on the actin filament. Troponin takes up ca++ ions from the sarcoplasm to initiate muscle contraction. In muscle troponin and tropomyosin combines to form troponin- tropomyosin system.

Sources of chemical energy

Adenosine triphosphates (ATP) - ATP is the immediate source of energy for muscle contraction. The breakdown of phosphate bond  of ATP releases maximum energy.
Anaerobic glycolysis:
Glucose -à 2 moles of lactic acid +8ATPs.
Aerobic glycolysis coupled with Kreb‘s cycle:
Glucose --à6 CO2 + 6H2O +38 ATPs.
Creatine phosphate (CP) or phosphagen - It forms a reservoir of high energy phosphate in the muscle.It cannot be used as a direct source of energy. It can be used for regeneration of ATP from ADP.
Creatine phosphate----------àcreatine + phosphoric acid
Phosphoric acid +ADP -------à ATP
Glucose – glycolysis as a source of energy - Glucose is stored in the muscle in the form of glycogen. Muscle glycogen is converted into glucose by glycogenolysis. Glucose is oxidized by glycolysis.
C6H12O6   + 6O2--------à6CO2 +6H2O +38 ATP

Cori’s Lactic acid cycle

The oxidation of lactic acid to carbon dioxide produces energy for the reconversion of ADP to ATP. The lactic acid produced in the muscle contraction passes into blood stream and is transported to the liver. Within the liver, lactic acid is converted to liver glycogen and then to blood glucose. The conversion of lactic acid to glycogen requires oxygen. Muscle glycogen comes only from the glucose of the blood.
Biochemical basis of muscle stimulation
The stimulation of nerve from the central nervous system initiates electrical changes in the muscle. The depolarization of sarcolemma is caused by the sudden influx of Na+ ions and efflux of k+ ions. The nerve impulse spreads in the muscle and releases Ca2+ ions from the sarcoplasmic reticulum. The flooding of Ca2+ ions starts the contractile machinery.
       Molecular changes during muscle contraction
The Ca2+ ions bind to the troponin molecules. Troponin – Ca2+ complex removes tropomyosin blockage of actin sites. The heads of myosin – ATP complex form Cross-bridges to actin filament. The hydrolysis of ATP induces conformational changes in the heads of myosin.
1 mole of Actin + 3 moles of myosin à Actomyosin
       Molecular changes during muscle relaxation
The Ca2+ ions are sequestered from actin filament by sacroplasmic reticulum. The Ca2+ ions are released from troponin – Ca2+ complex. Troponin permits tropomyosin return to blocking position. Then there is a separation of myosin-actin cross-bridges. ATP – myosin Complex reformed in heads of thick filament.
Physical changes during muscle contraction
Heat production -liberation of heat is always associated with muscle contraction.
Electricity generation -small amount of electrical energy is released.
Volume changes –negligible changes in volume occur.
Change in optical properties -changes in the birefringence and transparency occur at the muscle fiber.
Sound production -muscle sound noted during contraction.
Theories/ models  of muscle contraction
Sliding filament theory 
This theory was evolved independently and more or less simultaneously by A.F Huxley and H.E. Huxley around 1950s. According to this theory, the force of contraction is developed by the cross bridges in the overlap region. The active shortening is caused by the movement of the cross bridges, which causes one filament to slide over the other. During muscle contraction, the actin filaments alone show movement. But the myosin filaments remain static. The mechanical movement utilizes the energy derived from the breakdown of ATP molecules.
 Electrochemical theory – Davies model (1963)
 In the resting state, the cross        bridges are in slanting position due to negative charges in both the basal and tips of cross bridges due to the concentration of magnesium ions. After the stimulation of muscle, the release of calcium ions change the electrical character which leads to mutual repulsion and shortening of the cross bridges. The attachment of cross bridges on the actin filaments cause sliding of actin filaments while myosin filaments remain static.
Rowboat model- Huxley-Simmons model – According to this scheme, there are flexible hinges on myosin: one between S1 heads and the long rods and the second between S2 and the LMM at the trypsin reaction site. When the head piece (S1) binds to the exposed site on actin it is thought to rotate. This type of rotation occurs simultaneously at numerous locations of actin – myosin filaments which cause shortening of the muscle. The energy for this process derived from the hydrolysis of ATP.

Friday, February 28, 2014

Ecological Niche Concept

                                         Ecological niche is a term used to denote the position of a species within an ecosystem. The term includes all of the interactions between a species and the biotic and abiotic environments. It is the functional role/position of an organism within an ecological community. The term ecological niche was coined by the naturalist Joseph Grinnell in 1917.
                                        Ecological niche is the total way of life of a species or functional role (its profession) in an ecosystem. It involves all the physical, chemical and biological conditions. Niche focuses on the environmental factors that influence growth, survival and reproduction of a species.
                                       At present an ecological niche is characterized in relation to 3 principal axes that combine most of the relevant variables of the environment: a habitat axis (climatic and physico-chemical variables), a trophic axis (feeding relationships) and a temporal axis (mode of use of food resources and of land cover as a function of time).


Definition of niche concept

Charles Elton (1927) defined niche as the status of an organism in its community to indicate what is doing and not merely what it looks like.
Hutchinson (1944-58) defined niche as the sum of all the environmental factors acting on the organism.
Odum (1971) defined the niche of a species as its role in the ecosystem. He referred the habitat as address of the species and niche as the profession of a species.
Difference between habitat and niche
Habitat is the living place of an organism. In other words habitat is a suitable environment or habitable place for a species.
But niche is the role of a species in a community and also its relationship with its environment. Niche is the ecological function of a species.

Objective of niche concept

The niche concept was formulated to determine the role of organisms in a community (Hutchinson, 1957).

Types of niches

Fundamental niche – a species’ niche is usually less extensive, when competitors and predators present. Its niche in the absence of these organisms is known as its fundamental niche.
Realized niche – It is the portion of the fundamental niche that a species actually occupies.
Functional niche – refers to a species position in food webs and trophic chains.
Spatial niche – the niche occupied by a species is favourable to it because it furnishes a suitable substratum and microclimate to it.
Trophic niche – two species living in the same habitat, may have different food habits. For example elephant, rabbit and deer are three types of herbivores living in a forest habitat. But each of them occupies different trophic niches. Elephant feeds on large trees; deer feeds on shrubs and rabbit feeds on herbs and grasses.
Ecological niche - the position of a species in a community.
Food niche - the food choice of a species in a trophic level.
Time niche -animals tend to be active at different times of the day e.g.,diurnal/nocturnal.
Place niche -different species foraging in different ways and in different places.

Joseph Grinnell’s concept (1917) of niche as habitat

Grinnell referred niche as an organism’s physical environment. It is the ultimate distributional unit. No two species in the same general territory can occupy the same ecological niche for long time. Grinnell’s niche is now referred as spatial niche.

Eltonian’s concept (1927) of niche as functional unit

Charles Elton defined the niche as the ecological role the organism plays in the community. He considered niche as its occupation. It refers to how an organism transforms energy, behaves, responds to and modifies its physical and biotic environment.

Hutchinson’s concept (1957) of niche as multidimensional hypervolume space

Hutchinson suggested that niche could be modelled as an imaginary space with many dimensions. Each dimension or axis represents the range of an environmental condition or resource that is required by the species i.e. abiotic or environmental factors axis, food resource axis and function of time axis.
Niche overlap – the degree to which two species share niche space. This is assumed to be proportional to the degree of competition. In the absence of competition, niches do not overlap. But under intense competition, there is greater overlap.
Niche breadth – it is a product of niche overlap. The narrower the niche, the more specialized the species.
Niche separation – It is the distance between the mean resource use curves for two species. It may be close or far.

Significance of ecological niches

1.    By occupying different spatial or trophic niches, animals escape from competition.
2.    The niche occupied by a species is favourable to it because it contributes a suitable substratum and microclimate.
3.    The segregation of each species into different niches permits the full exploitation of the available resources.
4.    The segregation into niches avoids confusion of activities between organisms in the community. It gives more orderly and efficient life patterns for each species.

5.    Overlapping niches leads to competition between two species and in the competition one species will be replaced by another – competitive exclusion principle.

Saturday, February 22, 2014

Natural resource management

Introduction
Natural resources are derived from the environment. People depend on natural resources for their survival. Natural resources can be the land, the soil, the water, the air, the plants and the animals. All life on earth depends on four vital natural resources such as air, water, soil and sunlight.  Some resources that can be replaced after use by natural processes are called renewable. The common examples include air, water, soil, living organisms and sunlight.  Natural resources that cannot be replaced are termed non-renewable. Fossil fuels are an example of non-renewable natural resources. Non-renewable resources exist in a fixed amount and can be replaced by processes that take millions of years. Extracting, processing and using natural resources create pollution of air, water and land. But sustainable conservation is using natural resources wisely and not contributing to any environmental pollution.



Definition of natural resource

A resource is a form of matter or energy which is essential for the functioning of organisms, populations and ecosystems.
In other words, natural resources are the materials or substances obtained from nature which are required for the survival and prosperity of human beings.

Types of natural resources

Natural resources can be classified on the basis of their origin as abiotic and biotic resources.
Abiotic resources are derived from the non-living world. e.g. land, water, air.
Biotic resources are derived from living organisms. E.g. fish, forests.
Natural resources are mostly classified on the basis of their continual utility as renewable and non-renewable resources.
Renewable natural resources – Renewable resources are natural resources that can be replenished naturally in a short period of time. These resources are inexhaustible in supply. These resources have the ability to regenerate within a few years or decades. Renewable resources are vulnerable to human abuse. E.g. freshwater, oxygen, trees and fish. Renewable natural resources can be either organic (e.g. plant and animal species) or inorganic renewable natural resources (e.g. water, air)
Non-renewable natural resources – A non-renewable resource is a natural resource that cannot be re-made or re-grow at a scale comparable to its consumption. They are fixed in quantity and can be depleted. They take very long time to form. The replenishment time may range from 100s to million years.  The extraction of non-renewable resources always damages the environment. Coal, oil and natural gas are considered non-renewable because they take million of years to form. They are the product of the fossilized remains of dead plants and animals that have been exposed to heat and pressure deep within the earth’s crust. Hence these resources are called fossil fuels. E.g. metals, minerals and fossil fuels.
Non-renewable natural resources can be subdivided into recyclable and non-recyclable resources.
Recyclable natural resources are those that can be collected after their use and can be recycled for future requirements. E.g. metals.
Some natural resources are called non-recyclable because they cannot be recycled after their use. E.g.fossil fuels, natural energy.
All natural resources can also be classified on the basis of their natural amounts of availability into two categories:
1.    Inexhaustible natural resources
2.     Exhaustible natural resources
Inexhaustible natural resources are available in huge amounts in the nature and are replenished after their consumption.  E.g. solar energy, wind power, rain fall, tidal power and hydropower.
Exhaustible natural resources are available in limited amounts in the nature and are not replenished at the same rate as consumption. E.g. fossil fuels, metals and minerals.
Natural resources can also be classified into potential and actual natural resources based on their stage of development.
1.    Potential natural resources
2.     Actual natural resources
Potential natural resources – are the resources which are found in a region whose quality and quantity have not been determined for utilization. They can be used only in the future. E.g.  Distribution of mineral oil wealth in India.
Actual natural resources – are the resources whose quality and quantity have been determined for utilization and are being used in the present time. E.g. availability of oil and natural gas from Mumbai offshore region.
Flow natural resources – It does not remain in one location and moves about because of natural actions in the physical environment. It is replaced by natural processes whether humans use them or not. The quantity is expressed in annual rates. E.g. solar energy, freshwater, waves and tides.
Stock natural resources – It can be permanently expended and whose quantity is usually expressed in absolute amounts. E.g. coal and petroleum deposits.

Forest resources

Forests are a highly valuable and most abundant resource. They are diverse in composition and density. Forests cover about 30% of the earth’s surface. They are inhabited by a variety life forms.  Forests have warm climate with adequate rainfall.  Forests have tall and dense trees with many wild animals. The soil of the forests is rich in organic matter and nutrients. Forests play a major role in the water cycle, which includes circulation, transformation and replenishment of freshwater from all of earth’s ecosystems. Forests are essential for the conservation of biodiversity, water and soil resources. Forests contribute substantially to the economic development of a country. Forests are usually defined as land areas dominated by trees, where the tree canopy covers at least 10% of the ground area.  A healthy forest is composed of many more unique ecosystems. Over two – thirds of known land-based species live in forests.  In other words forests are home to 50-90% of earth’s species. In many countries, forests are the primary targets for agricultural and urban expansion.
Importance of forests
Forests are an important renewable natural resource.  Forests are the natural wealth of a country. They are the store houses of biodiversity. Forests provide various goods and services and also maintain the life-support systems of the  earth. Forests provide approximately 1.6 billion people with food, medicines, fuel and other basic necessities. Forests have the following three types of functions:
1.    Productive functions – Forests yield timber, bamboos, food and a variety of products such as alkaloids, essential oils, resins, latex, rubber and medicines.
2.    Protective functions – Forests conserve soil and water. They prevent drought and protect against wind, cold, heat, radiation and noise.
3.    Regulative functions – Forests absorb, store and release gases, water, minerals and sun light. They regulate floods, droughts and biogeochemical cycles.
Deforestation
Deforestation is the direct human-induced conversion of forest to non-forest land use such as agriculture, grazing or urban/industrial development. Deforestation is defined as the destruction of forest due to over-cutting of trees.
Effects of deforestation
Deforestation causes irreversible environmental damage. Deforestation accelerates extinction of plants,  animals and microbes. Deforestation disrupts cloud formation and rainfall. Deforestation threatens the livelihoods of tribal communities.
Deforestation increases soil erosion and decreases soil fertility. Deforestation increases greenhouse gases and warming of the environment.
Conservation and management of forests
An increase in forest area is initiated by  the following ways:
1.    Afforestation programmes- planting of trees on land that was not previously forested.
2.    Reforestation programmes- re-establishment of forest formation.
3.    Conservation of reserve forests - forests are protected in National parks, Sanctuaries, Sacred Grooves and Biosphere reserves.
4.    Chipko movement – This tree hugging movement was first started in Tehri Garhwal district in Uttarkhand in 1972. The movement was popularized by Sunderlal Bahuguna . Under this campaign, people started hugging trees whenever forest contractors tried to cut the trees.
5.    Production or commercial forestry – includes three programmes i.e. social forestry, agro forestry and urban forestry.

Mineral resources

Minerals are a major form of non-renewable resource. A rock is an aggregate of minerals. Minerals are formed over a period of millions of years in the earth’s crust. Mineral are naturally occurring inorganic compounds of definite chemical composition and characteristic physical properties.
A mineral can be defined as a pure element or chemical compound that is normally crystalline and that has been formed as a result of geological processes.
 There are more than 3,500 known minerals. Minerals range in composition from pure elements and simple salts to very complex silicates. The study of minerals is called mineralogy. Global distribution of minerals is not uniform. Minerals can be grouped into two broad categories:
1.    Metallic minerals – These mineral are found in the form of ores. The metal ores contain several impurities. E.g. iron, copper, gold and lead.
2.    Non-metallic minerals – these minerals are composed of elements like silicon, oxygen, calcium, magnesium, iron, aluminium etc. E.g  quartz, graphite, feldspar, dolomite, calcite, laterite, diamonds, sulphur, potash, salts, coal and petroleum.
Conservation of mineral resources
1.    Recycling – minerals in products can be recycled.
2.    Reuse – the reuse products more economical
3.    Substitution – scarce minerals can be substituted with more abundant minerals.
4.    Decreased consumption – consumers can decrease their mineral consumption.
5.    Use of industrial waste – one industry may use the waste products of another industry.

Water resources

Water is a renewable and limiting resource. It covers approximately 71% of the planet earth and constitutes 60-70 weight % of the living world. About 97.5 percent of the water on earth is strongly saline (sea water) the remaining 2.5 percent is freshwater. Pure, usable freshwater is only 0.3 percent.  Freshwater is therefore, a treasure to protect and conserve.  Water is a unique molecule because it is the only natural substance that exists in three physical states like liquid, solid (ice) and gas (steam) on earth.   Water is often referred to as a universal solvent because it has an unusual ability to dissolve other chemical substances. Water is the cellular medium within which all biochemical reactions can occur in living organisms. It is the transport medium for food, oxygen and other things needed by cells. Water is one of the vital elements for human life. The total amount of freshwater on earth is 1.4 billion cubic kilometres, but only around 41000 km3 circulates through hydrologic cycle. All the freshwater on earth comes from the rainfall. The amount of rainfall that falls on the continents is finite.
Sources of freshwater resources
Surface water – the freshwater exists in surface water bodies like streams, lakes, ponds, wetlands and man-made reservoirs. It is the most important resource of freshwater available for agricullure, industrial and domestic uses. The surface water flows in our country through 14 major rivers viz. Indus, Ganga, Brahmaputra, Godavari, Mahanadi, Krishna, Cauveri, Tapti, Mahi etc.
Ground water - the freshwater, which is received through rainfall, infiltrates into the soil and supplements underground water reserves. The hydrologic data suggests that the annual rainfall in India is about 400 m ha m. According to an estimate about 9.86% of the total freshwater resources are in the form of ground water. Of all freshwaters, ground water is less likely to be get polluted.
Integrated water resources management (IWRM)
IWRM has been defined as “a coordinated, goal-directed process for controlling the development and use of river, lake, ocean, wetland and other water assets”. It is a comprehensive approach for implementing efficient, equitable and sustainable solution to water and development problems.
Sustainable water management
1.    Construct dams and many small reservoirs.
2.    Develop small catchment dams and protect wetlands.
3.    Desalinize sea water into usable water.
4.    Divert freshwater canals/rivers to dry areas.
5.    Dredge and desilt rivers and water bodies regularly.

Food resources

Food is one of the basic needs of every living organism. Good food is indispensable for health and well being of humans at all stages of life. The food is composed of organic molecules such as carbohydrates, proteins and fats. The body uses some of the molecules in food to function correctly and stay healthy. The key to good health lies in maintaining balanced nutrition. The major source of human food includes plants or plant products and animal products. The plant products include cereals, pulses, vegetables fruits and spices. The animal products constitute milk or milk products, eggs and meat. Agricultural crops  constitute an important food resource. About 300 crops are grown for food and only 100 are used on a large scale.
Of the earth’s 30,000 plants with edible parts, we eat only about 30 plants. Of this only 15 plants and 8 animal species supply 90% of our food. Four crops such as wheat, rice, corn and potato make up more of the world’s total food production than all others combined. Grain provides about half the world’s calories with 2/3 people eat mainly vegetarian diet. As incomes rise, people consume more animal proteins in the form of meat, eggs, milk, cheese and other products of domesticated live stock. Fruits and vegetables also make a large contribution to human diet. They contain high levels of vitamins, minerals, dietary fibre, and complex carbohydrates. Domesticated animals form an important food source. The major domesticated animals used as food are ruminants such as cattle, sheep, goats, camel, reindeer etc.
Negative Impact of modern agriculture
1.    Declining genetic diversity – leads to loss of genetic diversity of indigenous crop varieties. Selective  farming   causes unformity of crops, monocultures and disease outbreaks.
2.    Soil erosion – Clearing  land for farming may result in soil erosion.
3.    Salinization of soil – soil builds up salt concentration due water logging. Intensive farming reduces soil fertility and productivity.
4.    Desertification – over-grazing is responsible for desertification.
5.    Fertilizer pollution – excessive accumulation of nitrates and phosphates is soil. Rainwater run off cause eutrophication of  surface water bodies and contamination of ground water.
6.    Pesticide pollution – over use of herbicides and insecticides cause chemical contamination of soil; pesticides can enter food chains; pesticides kill beneficial soil organisms.
7.    Other impacts – farming takes up land and reducing habitats and wildlife; irrigation (watering of crops) may take too much of water from rivers and deprive downstream habitats of water.
Sustainable Agriculture
A sustainable agriculture must be economically viable, socially responsible and ecologically sound. An agriculture that uses up or degrade its natural resource base or pollutes the natural environment eventually will lose its ability to produce. Such agriculture is not sustainable.
World food problems
In 1960s, the Green Revolution and advances in technology greatly increased the food production in many countries. There is a saying that “there is plenty of food on the planet, it is just a problem of distribution.” Global demand for food increased as a function of population growth for several decades. At one stage, the world human population surpassed the global food production. After 2001, the agriculture around the world was drastically affected by extreme weather conditions. Diseases affecting the crops and livestock have devastating effects on food production. By 2006, world cereal production has fallen by 2.1 percent. The prices for basic foods such as rice, wheat, and corn have risen 83% since 2005. Many countries experience perpetual food shortages. This results in widespread hunger and malnutrition.
Food security – refers to the availability of food and one’s access to it. Food security is the availability of enough food and access to all people at all times for an active and healthy life. The WHO defines three facets of food security:
1.    Food availability – availability of sufficient quantities of food on a consistent basis.
2.    Food access – sufficient household income to obtain appropriate foods.
3.    Food use – access to safe water, good sanitation and basic health care can make a difference in nutritional well being. They have an impact on body’s ability to utilize consumed foods.

Land resources

Land is one of the major life supporting systems. Land occupies 38% (one fifth) of the total earth’s space. Soil covers about four fifth of the land area. Soil is a biologically active, complex mixture of minerals, organic materials, living organisms, air and water. Soil is an interface between biosphere, lithosphere, hydrosphere and atmosphere. Soil is the outer most layer of the earth’s crust (Earth’s living skin-1/3 of surface). Soil is a vital resource that provides food, feed, fuel and fiber.
Soils have four major components: mineral matter, organic matter, air, and water. Mineral matter contains three fractions, sand, silt, and clay. Organic matter contains appreciable quantities of nitrogen, phosphorus and sulfur. Air and water occupy the pore spaces in soils. A fertile soil is a living soil containing billions of living organisms in every cubic centimetre. Plant nutrients in the soil are regularly regenerated by decay, decomposition and mineralization of organic matter by living organisms. Living communities in the soil are the basis for its fertility and other properties, which promote healthy plant life. Depletion of soil fertility is caused by removal of vegetation, water logging and application of non-degradable chemicals.
Sustainable Soil Conversion
1. Conservational tillage – ploughing improves soil permeability, soil moisture and nutrients.
2. Organic farming – More organic inputs to soil.
3. Crop rotation – growing legumes after cereal crops.
4. Contour ploughing – ploughing with alternate furrows and ridges.
5. Mulching – Soil is allowed to remain united and is covered with plant litter.
6. Strip cropping – planting in rows or strips.
7. Terrace farming – Hill slope is converted into terraces.
8.  Agrostological methods – Grasses are grown in rotation or along with agricultural crops.
9.  Afforestation – Trees or wind breaks are planted in deserts.

Air resources

Air is one of the natural resources and is a life-supporting system. Air is one of the basic needs of life. It forms the immediate environment of the earth and biosphere. The “air” is actually a mixture of gases. For example, 78% of the atmosphere is made of the nitrogen gas, 21% is composed of oxygen, and 0.9% is made up of argon. The atmosphere contains gases, suspended liquids, and solids that entirely surround the earth. The earth's gravity pulls these gases, liquids, and solids toward the surface.
The composition of air is not fixed. It changes according to the time of the day, the conditions of environment and humans’ activities. Air pollution refers to the presence of foreign particles in air which can affect the quantity of air and the health of living things. Carbon dioxide is supplied into the atmosphere by plant and animal respiration, the decay of organic material, volcanic eruptions, and natural and anthropogenic combustion. Carbon dioxide is removed from the atmosphere by photosynthesis. CO2 is an important greenhouse gas.
Uses of air resource
1.    Organisms can respire oxygen and survive in the world.
2.    Rainfall in any area depends on wind or air current.
3.    Wind has been used to generate electricity.
4.    Ozone protects the earth from uv rays.
5.    Nitrogen is an important mineral nutrients for plants.
6.    Air acts as a medium for some microorganisms.
7.    Water vapour maintains atmospheric humidity and in precipitation.

Energy resources

         Energy is the capacity to do work i.e.to move matter. Energy is our most important resource. Energy availability may limit economic and population growth. Most of our energy comes from fossil fuels. Sun is the primary source of energy. According to the first law of thermodynamics, ‘energy and matter can neither be created nor destroyed’. The second law defines that, ‘energy cannot be completely recycled’. Life depends on energy flow through ordered structures of matter.
Energy comes in different forms such as heat (thermal), light (radiant), mechanical, electrical, chemical and nuclear energy. There are two types of energy:
1.    Potential energy – stored energy
2.    Kinetic energy – working energy
Energy is produced usually from fuel burning and atom splitting.  Heat is considered to be low quality energy because it is dispersed. It is commonly a by-product of mechanical work. Electricity is a high quality form of energy because  it can be transported through wires, stored in batteries and used to make other forms of energy.

Kinds of energy resources

Energy resources can be classified into two groups:
1.    Primary energy resources
2.    Secondary energy resources
Primary energy is an energy form found in nature.  Secondary energy refers to all sources of energy those results from the transformation of primary energy resources. For example electricity is a secondary energy obtained from burning coal in a thermal power plant. Primary energy resources can be subdivided into two forms as:
       i.            Non-renewable energy resources
     ii.            Renewable energy resources
The energy sources can also be classified into conventional and non- conventional energy sources.
Conventional energy sources – energy sources which we are using it for a long time at a commercial scale. e.g. fossil fuels and hydroelectric power.
Non-conventional energy sources – energy sources which are in the process of development over the past few years. They are available in plenty and can be replenished naturally. E.g. solar, wind, tidal, geothermal and biomass.

Non-renewable energy resources

A non-renewable energy can be defined as a resource that cannot be reproduced or re-made at a scale comparable to its consumption. Once depleted, there will be no more available for future use. For example fossil fuels (coal, petroleum and natural gas) take thousands of years to form naturally and cannot be replenished in a short period of time.  Impurities in fossil fuels are a major source of pollution. The burning of fossil fuels produce large amounts of carbon dioxide, which contributes to global warming.

Renewable energy

Renewable energy resources can be defined as the resources that are naturally replenished, inexhaustible in duration but limited in the amount of energy that is available per unit of time. Renewable energy is obtained from natural resources such as sunlight, wind, tides, waves and geothermal heat. Common applications of renewable energies are electricity generation and motor fuels. Renewable energy resources currently contribute to approximately 18% of total global energy consumption. Examples include:
1.    solar energy ,
2.    hydro power,
3.    tidal power,
4.    wind  energy and
5.    geothermal (energy from the heat inside the earth).

Sustainable conservation of natural resources

Sustainability is a state of balance between resource use and the regenerative capacity of the earth. It can be described as ‘securing quality of life within the limits of nature’. The sustainability can be defined as “forms of progress that meet the needs of the present without compromising the ability of future generations to meet their needs”(Our Common Future 1987 : The World Commission on Environment and Development). Achieving sustainability is a balancing act between current needs and future needs.
Sustainability lies in the interplay of environmental quality, economic vitality and social equity. Ecological sustainability is the maintenance of life support systems and the achievement of natural extinction rate of species. Environmental sustainability refers to the maintenance of natural capital (e.g. natural resources).

The measures for sustainable conservation

Stabilizing population growth- The over-growth of human population has been a critical factor in over-utilizing the natural resources. Human race must willingly practice population control. Population should be stabilized at the level of carrying capacity of the planet earth.
Change-over in energy  use- A change-over is necessary from non-renewable polluting energy to renewable and non-polluting alternative energy resources.
Change – over in technology -A change-over is needed from resource intensive and population – prone technologies to environmental – friendly technologies.
Change–over in economy -The economic development must be environmentally sensitive and sustainable.
Scientific conservation - Conservation is the ‘gospel of efficiency’. Scientific methods should be followed to manage nature and natural resources efficiently.